Spring 2026 Subject: Ecology, Evolution and Environmental Biology

Spring 2026 Ecology, Evolution and Environmental Biology UN1011 BEHAVIOR BIOL-LIVING PRIMATES
Section 001	BEHAVIOR BIOL-LIVING PRIM Call Number: 13083 Points: 3 View in Vergil Day/Time: MW 1:10pm-2:25pm Location: To be announced Enrollment: 0 students (50 max) as of October 13, 2025 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN1111 BEHAVIORAL BIOL-DISC
Section 001	Call Number: 13086 Points: 0 View in Vergil Day/Time: W 6:10pm-7:00pm Location: To be announced Enrollment: 0 students (25 max) as of October 13, 2025 Instructor: Marina Cords
Section 002	Call Number: 13091 Points: 0 View in Vergil Day/Time: F 11:10am-12:00pm Location: To be announced Enrollment: 0 students (25 max) as of October 13, 2025 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN2002 ENVIRONMENTAL BIOLOGY II
Section 001	Call Number: 13096 Points: 4 View in Vergil Day/Time: MW 11:40am-12:55pm Location: To be announced Enrollment: 0 students (40 max) as of October 13, 2025 Notes: Labs meets W or Th 4:10-7:00. Students choose a lab section Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology UN3087 CONSERVATION BIOLOGY
Section 001	Call Number: 13098 Points: 3 View in Vergil Day/Time: M 6:10pm-8:00pm Location: To be announced Enrollment: 0 students (30 max) as of October 13, 2025 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3187 CONSERVATION BIOLOGY-DISC
Section 001	Call Number: 13104 Points: 0 View in Vergil Day/Time: W 6:10pm-7:00pm Location: To be announced Enrollment: 0 students (20 max) as of October 13, 2025 Instructor: Dustin Partridge
Section 002	Call Number: 13105 Points: 0 View in Vergil Day/Time: R 6:10pm-7:00pm Location: To be announced Enrollment: 0 students (20 max) as of October 13, 2025 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3215 FORENSIC OSTEOLOGY
Section 001	Call Number: 13163 Points: 3 View in Vergil Day/Time: MW 4:10pm-6:00pm Location: To be announced Enrollment: 0 students (15 max) as of October 13, 2025 Notes: EBHS students have priority at first class session Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3910 THE NEANDERTALS
Section 001	Call Number: 13168 Points: 4 View in Vergil Day/Time: T 2:10pm-4:00pm Location: To be announced Enrollment: 0 students (12 max) as of October 13, 2025 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3970 Biological Basis of Human Variation
Section 001	Bio Basis Human Variation Call Number: 13170 Points: 4 View in Vergil Day/Time: TR 11:40am-12:55pm Location: To be announced Enrollment: 0 students (18 max) as of October 13, 2025 Instructor: Volney K Friedrich
Spring 2026 Ecology, Evolution and Environmental Biology UN3992 THESIS RESEARCH SEMINAR
Section 001	Call Number: 13188 Points: 3 View in Vergil Day/Time: R 4:10pm-6:00pm Location: To be announced Enrollment: 0 students (20 max) as of October 13, 2025 Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology UN3994 EBHS SENIOR THESIS SEMINAR
Section 001	EBHS SENIOR THESIS SEMINA Call Number: 13195 Points: 3 View in Vergil Day/Time: M 1:10pm-3:00pm Location: To be announced Enrollment: 0 students (6 max) as of October 13, 2025 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3998 INDEPENDENT STUDY
Section 001	Call Number: 13206 Points: 1-3 View in Vergil Enrollment: 0 students (6 max) as of October 13, 2025 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology GU4015 ANIMAL COMMUN:PRIMATE PERSP
Section 001	ANIMAL COMMUN:PRIMATE PER Call Number: 13214 Points: 3 View in Vergil Day/Time: TR 10:10am-11:25am Location: To be announced Enrollment: 0 students (15 max) as of October 13, 2025 Instructor: Alba Lucia Morales Jimenez
Spring 2026 Ecology, Evolution and Environmental Biology GU4055 PRINCIPLES & APPS MOD DNA SEQUENCING TEC
Section 001	PRINCIPLES & APPS MOD DNA Call Number: 13238 Points: 3 View in Vergil Day/Time: MW 1:10pm-2:25pm Location: To be announced Enrollment: 0 students (15 max) as of October 13, 2025 Notes: Introductory biology and/or genetics. Computer programming e Instructor: Andres Bendesky
Spring 2026 Ecology, Evolution and Environmental Biology GU4105 Intermediate Statistics for Ecology and
Section 001	Intermed Stats-Ecology & Call Number: 13242 Points: 3 View in Vergil Day/Time: M 6:10pm-8:00pm Location: To be announced Day/Time: T 6:10pm-7:00pm Location: To be announced Enrollment: 0 students (20 max) as of October 13, 2025 Notes: Some prior intro course to R and Stats required, or equivale Instructor: Steffen Foerster
Spring 2026 Ecology, Evolution and Environmental Biology GU4112 ICHTHYOLOGY
Section 001	Call Number: 13243 Points: 3 View in Vergil Day/Time: F 10:10am-12:00pm Location: To be announced Enrollment: 0 students (15 max) as of October 13, 2025
Spring 2026 Ecology, Evolution and Environmental Biology GU4126 INTRO TO CONSERVATION GENETICS
Section 001	INTRO TO CONSERVATION GEN Call Number: 13245 Points: 3 View in Vergil Day/Time: R 10:10am-12:00pm Location: To be announced Enrollment: 0 students (15 max) as of October 13, 2025 Instructor: Rachel S Welt
Spring 2026 Ecology, Evolution and Environmental Biology GU4135 URBAN ECOLOGY & DESIGN
Section 001	Call Number: 13247 Points: 3 View in Vergil Day/Time: T 12:10pm-2:00pm Location: To be announced Enrollment: 0 students (40 max) as of October 13, 2025 Notes: Some weeks will have Friday field trips Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology GU4192 INTRO TO LANDSCAPE ANALYSIS
Section 001	INTRO TO LANDSCAPE ANALYS Call Number: 13250 Points: 3 View in Vergil Day/Time: M 8:40am-11:25am Location: To be announced Enrollment: 0 students (25 max) as of October 13, 2025
Spring 2026 Ecology, Evolution and Environmental Biology GU4201 ECO, BEHAVIOR & CONSERVATION OF MAMMALS
Section 001	ECO, BEHAVIOR & CONSERVAT Call Number: 13254 Points: 3 View in Vergil Day/Time: R 6:10pm-8:00pm Location: To be announced Enrollment: 0 students (12 max) as of October 13, 2025 Instructor: Scott Lapoint
Spring 2026 Ecology, Evolution and Environmental Biology GU4650 BIODIVERSITY & ECOSYSTEM PROC
Section 001	BIODIVERSITY & ECOSYSTEM Call Number: 13256 Points: 3 View in Vergil Day/Time: M 4:10pm-6:00pm Location: To be announced Enrollment: 0 students (25 max) as of October 13, 2025 Instructor: Shahid Naeem
Spring 2026 Ecology, Evolution and Environmental Biology GU4670 Introduction to Geographical Information
Section 001	Introduction to GIS Call Number: 13258 Points: 3 View in Vergil Day/Time: R 12:10pm-2:00pm Location: To be announced Enrollment: 0 students (30 max) as of October 13, 2025 Instructor: Viorel Popescu
Spring 2026 Ecology, Evolution and Environmental Biology GR5851 THESIS DEVELOPMENT SEMINAR
Section 001	THESIS DEVELOPMENT SEMINA Call Number: 13259 Points: 3 View in Vergil Day/Time: W 10:10am-12:00pm Location: To be announced Enrollment: 0 students (20 max) as of October 13, 2025 Instructor: Bekka Brodie
Spring 2026 Ecology, Evolution and Environmental Biology GR6110 FUNDAMENTALS OF EVOLUTION
Section 001	Call Number: 13260 Points: 3 View in Vergil Day/Time: TR 8:10am-10:00am Location: To be announced Enrollment: 0 students (30 max) as of October 13, 2025 Instructor: Kira Delmore
Spring 2026 Ecology, Evolution and Environmental Biology GR6300 RESEARCH SEMINAR
Section 001	Call Number: 13261 Points: 1 View in Vergil Day/Time: T 4:00pm-5:00pm Location: To be announced Day/Time: R 2:30pm-3:30pm Location: To be announced Enrollment: 0 students (100 max) as of October 13, 2025 Instructor: Maria Diuk-Wasser
Spring 2026 Ecology, Evolution and Environmental Biology GR6905 GRAD SEM IN CONSERVATION BIOL
Section 001	GRAD SEM IN CONSERVATION Call Number: 13262 Points: 3 View in Vergil Day/Time: T 10:10am-12:00pm Location: To be announced Enrollment: 0 students (25 max) as of October 13, 2025 Instructor: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology UN1011
BEHAVIOR BIOL-LIVING PRIMATES

Section 001

Call Number:: 13083
Points:: 3
Day/Time:: MW 1:10pm-2:25pm
Location:: To be announced
Enrollment:: 0 students (50 max) as of October 13, 2025
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN1111
BEHAVIORAL BIOL-DISC

Section 001

Call Number:: 13086
Points:: 0
Day/Time:: W 6:10pm-7:00pm
Location:: To be announced
Enrollment:: 0 students (25 max) as of October 13, 2025
Instructor:: Marina Cords

Section 002

Call Number:: 13091
Points:: 0
Day/Time:: F 11:10am-12:00pm
Location:: To be announced
Enrollment:: 0 students (25 max) as of October 13, 2025
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN2002
ENVIRONMENTAL BIOLOGY II

Section 001

Call Number:: 13096
Points:: 4
Day/Time:: MW 11:40am-12:55pm
Location:: To be announced
Enrollment:: 0 students (40 max) as of October 13, 2025
Notes:: Labs meets W or Th 4:10-7:00. Students choose a lab section
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology UN3087
CONSERVATION BIOLOGY

Section 001

Call Number:: 13098
Points:: 3
Day/Time:: M 6:10pm-8:00pm
Location:: To be announced
Enrollment:: 0 students (30 max) as of October 13, 2025
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3187
CONSERVATION BIOLOGY-DISC

Section 001

Call Number:: 13104
Points:: 0
Day/Time:: W 6:10pm-7:00pm
Location:: To be announced
Enrollment:: 0 students (20 max) as of October 13, 2025
Instructor:: Dustin Partridge

Section 002

Call Number:: 13105
Points:: 0
Day/Time:: R 6:10pm-7:00pm
Location:: To be announced
Enrollment:: 0 students (20 max) as of October 13, 2025
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3215
FORENSIC OSTEOLOGY

Section 001

Call Number:: 13163
Points:: 3
Day/Time:: MW 4:10pm-6:00pm
Location:: To be announced
Enrollment:: 0 students (15 max) as of October 13, 2025
Notes:: EBHS students have priority at first class session
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3910
THE NEANDERTALS

Section 001

Call Number:: 13168
Points:: 4
Day/Time:: T 2:10pm-4:00pm
Location:: To be announced
Enrollment:: 0 students (12 max) as of October 13, 2025
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3970
Biological Basis of Human Variation

Section 001

Call Number:: 13170
Points:: 4
Day/Time:: TR 11:40am-12:55pm
Location:: To be announced
Enrollment:: 0 students (18 max) as of October 13, 2025
Instructor:: Volney K Friedrich

Spring 2026 Ecology, Evolution and Environmental Biology UN3992
THESIS RESEARCH SEMINAR

Section 001

Call Number:: 13188
Points:: 3
Day/Time:: R 4:10pm-6:00pm
Location:: To be announced
Enrollment:: 0 students (20 max) as of October 13, 2025
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology UN3994
EBHS SENIOR THESIS SEMINAR

Section 001

Call Number:: 13195
Points:: 3
Day/Time:: M 1:10pm-3:00pm
Location:: To be announced
Enrollment:: 0 students (6 max) as of October 13, 2025
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3998
INDEPENDENT STUDY

Section 001

Call Number:: 13206
Points:: 1-3
Enrollment:: 0 students (6 max) as of October 13, 2025
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology GU4015
ANIMAL COMMUN:PRIMATE PERSP

Section 001

Call Number:: 13214
Points:: 3
Day/Time:: TR 10:10am-11:25am
Location:: To be announced
Enrollment:: 0 students (15 max) as of October 13, 2025
Instructor:: Alba Lucia Morales Jimenez

Spring 2026 Ecology, Evolution and Environmental Biology GU4055
PRINCIPLES & APPS MOD DNA SEQUENCING TEC

Section 001

Call Number:: 13238
Points:: 3
Day/Time:: MW 1:10pm-2:25pm
Location:: To be announced
Enrollment:: 0 students (15 max) as of October 13, 2025
Notes:: Introductory biology and/or genetics. Computer programming e
Instructor:: Andres Bendesky

Spring 2026 Ecology, Evolution and Environmental Biology GU4105
Intermediate Statistics for Ecology and

Section 001

Call Number:: 13242
Points:: 3
Day/Time:: M 6:10pm-8:00pm
Location:: To be announced
Day/Time:: T 6:10pm-7:00pm
Location:: To be announced
Enrollment:: 0 students (20 max) as of October 13, 2025
Notes:: Some prior intro course to R and Stats required, or equivale
Instructor:: Steffen Foerster

Spring 2026 Ecology, Evolution and Environmental Biology GU4112
ICHTHYOLOGY

Section 001

Call Number:: 13243
Points:: 3
Day/Time:: F 10:10am-12:00pm
Location:: To be announced
Enrollment:: 0 students (15 max) as of October 13, 2025

Spring 2026 Ecology, Evolution and Environmental Biology GU4126
INTRO TO CONSERVATION GENETICS

Section 001

Call Number:: 13245
Points:: 3
Day/Time:: R 10:10am-12:00pm
Location:: To be announced
Enrollment:: 0 students (15 max) as of October 13, 2025
Instructor:: Rachel S Welt

Spring 2026 Ecology, Evolution and Environmental Biology GU4135
URBAN ECOLOGY & DESIGN

Section 001

Call Number:: 13247
Points:: 3
Day/Time:: T 12:10pm-2:00pm
Location:: To be announced
Enrollment:: 0 students (40 max) as of October 13, 2025
Notes:: Some weeks will have Friday field trips
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology GU4192
INTRO TO LANDSCAPE ANALYSIS

Section 001

Call Number:: 13250
Points:: 3
Day/Time:: M 8:40am-11:25am
Location:: To be announced
Enrollment:: 0 students (25 max) as of October 13, 2025

Spring 2026 Ecology, Evolution and Environmental Biology GU4201
ECO, BEHAVIOR & CONSERVATION OF MAMMALS

Section 001

Call Number:: 13254
Points:: 3
Day/Time:: R 6:10pm-8:00pm
Location:: To be announced
Enrollment:: 0 students (12 max) as of October 13, 2025
Instructor:: Scott Lapoint

Spring 2026 Ecology, Evolution and Environmental Biology GU4650
BIODIVERSITY & ECOSYSTEM PROC

Section 001

Call Number:: 13256
Points:: 3
Day/Time:: M 4:10pm-6:00pm
Location:: To be announced
Enrollment:: 0 students (25 max) as of October 13, 2025
Instructor:: Shahid Naeem

Spring 2026 Ecology, Evolution and Environmental Biology GU4670
Introduction to Geographical Information

Section 001

Call Number:: 13258
Points:: 3
Day/Time:: R 12:10pm-2:00pm
Location:: To be announced
Enrollment:: 0 students (30 max) as of October 13, 2025
Instructor:: Viorel Popescu

Spring 2026 Ecology, Evolution and Environmental Biology GR5851
THESIS DEVELOPMENT SEMINAR

Section 001

Call Number:: 13259
Points:: 3
Day/Time:: W 10:10am-12:00pm
Location:: To be announced
Enrollment:: 0 students (20 max) as of October 13, 2025
Instructor:: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology GR6110
FUNDAMENTALS OF EVOLUTION

Section 001

Call Number:: 13260
Points:: 3
Day/Time:: TR 8:10am-10:00am
Location:: To be announced
Enrollment:: 0 students (30 max) as of October 13, 2025
Instructor:: Kira Delmore

Spring 2026 Ecology, Evolution and Environmental Biology GR6300
RESEARCH SEMINAR

Section 001

Call Number:: 13261
Points:: 1
Day/Time:: T 4:00pm-5:00pm
Location:: To be announced
Day/Time:: R 2:30pm-3:30pm
Location:: To be announced
Enrollment:: 0 students (100 max) as of October 13, 2025
Instructor:: Maria Diuk-Wasser

Spring 2026 Ecology, Evolution and Environmental Biology GR6905
GRAD SEM IN CONSERVATION BIOL

Section 001

Call Number:: 13262
Points:: 3
Day/Time:: T 10:10am-12:00pm
Location:: To be announced
Enrollment:: 0 students (25 max) as of October 13, 2025
Instructor:: Bekka Brodie

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.

Spring 2026 Subject: Ecology, Evolution and Environmental Biology

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BEHAVIOR BIOL-LIVING PRIM

Bio Basis Human Variation

EBHS SENIOR THESIS SEMINA

ANIMAL COMMUN:PRIMATE PER

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

BIODIVERSITY & ECOSYSTEM

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.

Spring 2026 Subject: Ecology, Evolution and Environmental Biology

For easier printing select theplain text version of this subject listingand be sure to print using LANDSCAPE mode

BEHAVIOR BIOL-LIVING PRIM

Bio Basis Human Variation

EBHS SENIOR THESIS SEMINA

ANIMAL COMMUN:PRIMATE PER

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

BIODIVERSITY & ECOSYSTEM

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

For easier printing select the
plain text version of this subject listing
and be sure to print using LANDSCAPE mode