Spring 2026 Department: Ecology, Evolution, and Environmental Biology

Spring 2026 Ecology, Evolution and Environmental Biology UN1011 BEHAVIOR BIOL-LIVING PRIMATES
Section 001	BEHAVIOR BIOL-LIVING PRIM Call Number: 13083 Points: 3 View in Vergil Day/Time: MW 1:10pm-2:25pm Location: 516 Hamilton Hall Enrollment: 32 students (50 max) as of January 9, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN1111 BEHAVIORAL BIOL-DISC
Section 001	Call Number: 13086 Points: 0 View in Vergil Day/Time: W 6:10pm-7:00pm Location: 1102 International Affairs Building Enrollment: 8 students (25 max) as of January 9, 2026 Instructor: Marina Cords
Section 002	Call Number: 13091 Points: 0 View in Vergil Day/Time: F 11:10am-12:00pm Location: 411 Kent Hall Enrollment: 3 students (25 max) as of January 9, 2026 Instructor: Marina Cords
Spring 2026 Ecology, Evolution and Environmental Biology UN2002 ENVIRONMENTAL BIOLOGY II
Section 001	Call Number: 13096 Points: 4 View in Vergil Day/Time: MW 11:40am-12:55pm Location: 501 Northwest Corner Building Enrollment: 20 students (40 max) as of January 9, 2026 Notes: Labs meets W or Th 4:10-7:00. Students choose a lab section Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology UN3087 CONSERVATION BIOLOGY
Section 001	Call Number: 13098 Points: 3 View in Vergil Day/Time: M 6:10pm-8:00pm Location: 603 Schermerhorn Hall [SCH] Enrollment: 30 students (30 max) as of January 9, 2026 / Full Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3187 CONSERVATION BIOLOGY-DISC
Section 001	Call Number: 13104 Points: 0 View in Vergil Day/Time: W 6:10pm-7:00pm Location: 104 Knox Hall Enrollment: 7 students (20 max) as of January 9, 2026 Instructor: Dustin Partridge
Section 002	Call Number: 13105 Points: 0 View in Vergil Day/Time: R 6:10pm-7:00pm Location: 652 Schermerhorn Hall [SCH] Enrollment: 0 students (20 max) as of January 9, 2026 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology UN3215 FORENSIC OSTEOLOGY
Section 001	Call Number: 13163 Points: 3 View in Vergil Day/Time: MW 4:10pm-6:00pm Location: 506 Schermerhorn Hall [SCH] Enrollment: 17 students (15 max) as of January 9, 2026 / Full Notes: EBHS students have priority at first class session Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3320 Biological Interactions in a Changing Wo
Section 001	Bio Interactions-Changing Call Number: 16840 Points: 3 View in Vergil Day/Time: TR 2:40pm-3:55pm Location: 506 Schermerhorn Hall [SCH] Enrollment: 7 students (20 max) as of January 9, 2026 Notes: Pre-requisites: Environmental Biology I & II or approval by Instructor: Rachel O Cohen
Spring 2026 Ecology, Evolution and Environmental Biology UN3910 THE NEANDERTALS
Section 001	Call Number: 13168 Points: 4 View in Vergil Day/Time: T 2:10pm-4:00pm Location: 327 Uris Hall Enrollment: 10 students (12 max) as of January 9, 2026 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3992 THESIS RESEARCH SEMINAR
Section 001	Call Number: 13188 Points: 3 View in Vergil Day/Time: R 4:10pm-6:00pm Location: To be announced Enrollment: 9 students (20 max) as of January 9, 2026 Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology UN3994 EBHS SENIOR THESIS SEMINAR
Section 001	EBHS SENIOR THESIS SEMINA Call Number: 13195 Points: 3 View in Vergil Day/Time: T 5:30pm-7:30pm Location: 1020 Schermerhorn Hall [SCH] Enrollment: 2 students (6 max) as of January 9, 2026 Instructor: Jill S Shapiro
Spring 2026 Ecology, Evolution and Environmental Biology UN3998 INDEPENDENT STUDY
Section 001	Call Number: 13206 Points: 1-3 View in Vergil Enrollment: 0 students (6 max) as of January 9, 2026 Instructor: Jill S Shapiro
Section 002	Call Number: 18062 Points: 1-3 View in Vergil Enrollment: 1 student (2 max) as of January 9, 2026 Instructor: Dustin Partridge
Spring 2026 Ecology, Evolution and Environmental Biology GU4055 PRINCIPLES & APPS MOD DNA SEQUENCING TEC
Section 001	PRINCIPLES & APPS MOD DNA Call Number: 13238 Points: 3 View in Vergil Day/Time: MW 1:10pm-2:25pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 14 students (15 max) as of January 9, 2026 Notes: Introductory biology and/or genetics. Computer programming e Instructor: Andres Bendesky
Spring 2026 Ecology, Evolution and Environmental Biology GU4105 Intermediate Statistics for Ecology and
Section 001	Intermed Stats-Ecology & Call Number: 13242 Points: 3 View in Vergil Day/Time: M 6:10pm-8:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Day/Time: T 6:10pm-7:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 20 students (20 max) as of January 9, 2026 / Full Notes: Some prior intro course to R and Stats required, or equivale Instructor: Steffen Foerster
Spring 2026 Ecology, Evolution and Environmental Biology GU4112 ICHTHYOLOGY
Section 001	Call Number: 13243 Points: 3 View in Vergil Day/Time: F 10:10am-12:40pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 14 students (15 max) as of January 9, 2026 Notes: Some classes will be held at AMNH Instructor: Pedro Braganca
Spring 2026 Ecology, Evolution and Environmental Biology GU4126 INTRO TO CONSERVATION GENETICS
Section 001	INTRO TO CONSERVATION GEN Call Number: 13245 Points: 3 View in Vergil Day/Time: R 10:10am-12:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 8 students (15 max) as of January 9, 2026 Instructor: Rachel S Welt
Spring 2026 Ecology, Evolution and Environmental Biology GU4135 URBAN ECOLOGY & DESIGN
Section 001	Call Number: 13247 Points: 3 View in Vergil Day/Time: T 12:10pm-2:00pm Location: 303 Uris Hall Enrollment: 45 students (45 max) as of January 9, 2026 / Full Notes: Some weeks will have Friday field trips Instructor: Matthew I Palmer
Spring 2026 Ecology, Evolution and Environmental Biology GU4192 INTRO TO LANDSCAPE ANALYSIS
Section 001	INTRO TO LANDSCAPE ANALYS Call Number: 13250 Points: 3 View in Vergil Day/Time: M 8:40am-11:25am Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 20 students (25 max) as of January 9, 2026 Instructors: Marcia N Macedo and Jay M Schoen
Spring 2026 Ecology, Evolution and Environmental Biology GU4201 ECO, BEHAVIOR & CONSERVATION OF MAMMALS
Section 001	ECO, BEHAVIOR & CONSERVAT Call Number: 13254 Points: 3 View in Vergil Day/Time: R 6:10pm-8:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 12 students (12 max) as of January 9, 2026 / Full Instructor: Scott Lapoint
Spring 2026 Ecology, Evolution and Environmental Biology GU4340 HUMAN ADAPTATION
Section 001	Call Number: 16846 Points: 3 View in Vergil Day/Time: TR 11:40am-12:55pm Location: 608 Schermerhorn Hall [SCH] Enrollment: 13 students (18 max) as of January 9, 2026 Instructor: Volney K Friedrich
Spring 2026 Ecology, Evolution and Environmental Biology GU4670 Introduction to Geographical Information
Section 001	Introduction to GIS Call Number: 13258 Points: 3 View in Vergil Day/Time: R 12:10pm-2:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 36 students (34 max) as of January 9, 2026 / Full Instructor: Viorel Popescu
Spring 2026 Ecology, Evolution and Environmental Biology GR5851 THESIS DEVELOPMENT SEMINAR
Section 001	THESIS DEVELOPMENT SEMINA Call Number: 13259 Points: 3 View in Vergil Day/Time: W 10:10am-12:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 8 students (20 max) as of January 9, 2026 Instructor: Bekka Brodie
Spring 2026 Ecology, Evolution and Environmental Biology GR6110 FUNDAMENTALS OF EVOLUTION
Section 001	Call Number: 13260 Points: 3 View in Vergil Day/Time: TR 8:10am-10:00am Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 22 students (30 max) as of January 9, 2026 Instructor: Kira Delmore
Spring 2026 Ecology, Evolution and Environmental Biology GR6300 RESEARCH SEMINAR
Section 001	Call Number: 13261 Points: 1 View in Vergil Day/Time: T 4:00pm-5:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Day/Time: R 2:30pm-3:30pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 27 students (100 max) as of January 9, 2026 Instructor: Maria Diuk-Wasser
Spring 2026 Ecology, Evolution and Environmental Biology GR6905 GRAD SEM IN CONSERVATION BIOL
Section 001	GRAD SEM IN CONSERVATION Call Number: 13262 Points: 3 View in Vergil Day/Time: T 10:10am-12:00pm Location: 1015 EXT Schermerhorn Hall [SCH] Enrollment: 15 students (25 max) as of January 9, 2026 Instructor: Bekka Brodie
Spring 2026 Ecology, Evolution and Environmental Biology GR9509 DIRECTED READINGS
Section 001	Call Number: 18063 Points: 1-6 View in Vergil Enrollment: 1 student (2 max) as of January 9, 2026 Instructor: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN1011
BEHAVIOR BIOL-LIVING PRIMATES

Section 001

Call Number:: 13083
Points:: 3
Day/Time:: MW 1:10pm-2:25pm
Location:: 516 Hamilton Hall
Enrollment:: 32 students (50 max) as of January 9, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN1111
BEHAVIORAL BIOL-DISC

Section 001

Call Number:: 13086
Points:: 0
Day/Time:: W 6:10pm-7:00pm
Location:: 1102 International Affairs Building
Enrollment:: 8 students (25 max) as of January 9, 2026
Instructor:: Marina Cords

Section 002

Call Number:: 13091
Points:: 0
Day/Time:: F 11:10am-12:00pm
Location:: 411 Kent Hall
Enrollment:: 3 students (25 max) as of January 9, 2026
Instructor:: Marina Cords

Spring 2026 Ecology, Evolution and Environmental Biology UN2002
ENVIRONMENTAL BIOLOGY II

Section 001

Call Number:: 13096
Points:: 4
Day/Time:: MW 11:40am-12:55pm
Location:: 501 Northwest Corner Building
Enrollment:: 20 students (40 max) as of January 9, 2026
Notes:: Labs meets W or Th 4:10-7:00. Students choose a lab section
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology UN3087
CONSERVATION BIOLOGY

Section 001

Call Number:: 13098
Points:: 3
Day/Time:: M 6:10pm-8:00pm
Location:: 603 Schermerhorn Hall [SCH]
Enrollment:: 30 students (30 max) as of January 9, 2026 / Full
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3187
CONSERVATION BIOLOGY-DISC

Section 001

Call Number:: 13104
Points:: 0
Day/Time:: W 6:10pm-7:00pm
Location:: 104 Knox Hall
Enrollment:: 7 students (20 max) as of January 9, 2026
Instructor:: Dustin Partridge

Section 002

Call Number:: 13105
Points:: 0
Day/Time:: R 6:10pm-7:00pm
Location:: 652 Schermerhorn Hall [SCH]
Enrollment:: 0 students (20 max) as of January 9, 2026
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology UN3215
FORENSIC OSTEOLOGY

Section 001

Call Number:: 13163
Points:: 3
Day/Time:: MW 4:10pm-6:00pm
Location:: 506 Schermerhorn Hall [SCH]
Enrollment:: 17 students (15 max) as of January 9, 2026 / Full
Notes:: EBHS students have priority at first class session
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3320
Biological Interactions in a Changing Wo

Section 001

Call Number:: 16840
Points:: 3
Day/Time:: TR 2:40pm-3:55pm
Location:: 506 Schermerhorn Hall [SCH]
Enrollment:: 7 students (20 max) as of January 9, 2026
Notes:: Pre-requisites: Environmental Biology I & II or approval by
Instructor:: Rachel O Cohen

Spring 2026 Ecology, Evolution and Environmental Biology UN3910
THE NEANDERTALS

Section 001

Call Number:: 13168
Points:: 4
Day/Time:: T 2:10pm-4:00pm
Location:: 327 Uris Hall
Enrollment:: 10 students (12 max) as of January 9, 2026
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3992
THESIS RESEARCH SEMINAR

Section 001

Call Number:: 13188
Points:: 3
Day/Time:: R 4:10pm-6:00pm
Location:: To be announced
Enrollment:: 9 students (20 max) as of January 9, 2026
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology UN3994
EBHS SENIOR THESIS SEMINAR

Section 001

Call Number:: 13195
Points:: 3
Day/Time:: T 5:30pm-7:30pm
Location:: 1020 Schermerhorn Hall [SCH]
Enrollment:: 2 students (6 max) as of January 9, 2026
Instructor:: Jill S Shapiro

Spring 2026 Ecology, Evolution and Environmental Biology UN3998
INDEPENDENT STUDY

Section 001

Call Number:: 13206
Points:: 1-3
Enrollment:: 0 students (6 max) as of January 9, 2026
Instructor:: Jill S Shapiro

Section 002

Call Number:: 18062
Points:: 1-3
Enrollment:: 1 student (2 max) as of January 9, 2026
Instructor:: Dustin Partridge

Spring 2026 Ecology, Evolution and Environmental Biology GU4055
PRINCIPLES & APPS MOD DNA SEQUENCING TEC

Section 001

Call Number:: 13238
Points:: 3
Day/Time:: MW 1:10pm-2:25pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 14 students (15 max) as of January 9, 2026
Notes:: Introductory biology and/or genetics. Computer programming e
Instructor:: Andres Bendesky

Spring 2026 Ecology, Evolution and Environmental Biology GU4105
Intermediate Statistics for Ecology and

Section 001

Call Number:: 13242
Points:: 3
Day/Time:: M 6:10pm-8:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Day/Time:: T 6:10pm-7:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 20 students (20 max) as of January 9, 2026 / Full
Notes:: Some prior intro course to R and Stats required, or equivale
Instructor:: Steffen Foerster

Spring 2026 Ecology, Evolution and Environmental Biology GU4112
ICHTHYOLOGY

Section 001

Call Number:: 13243
Points:: 3
Day/Time:: F 10:10am-12:40pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 14 students (15 max) as of January 9, 2026
Notes:: Some classes will be held at AMNH
Instructor:: Pedro Braganca

Spring 2026 Ecology, Evolution and Environmental Biology GU4126
INTRO TO CONSERVATION GENETICS

Section 001

Call Number:: 13245
Points:: 3
Day/Time:: R 10:10am-12:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 8 students (15 max) as of January 9, 2026
Instructor:: Rachel S Welt

Spring 2026 Ecology, Evolution and Environmental Biology GU4135
URBAN ECOLOGY & DESIGN

Section 001

Call Number:: 13247
Points:: 3
Day/Time:: T 12:10pm-2:00pm
Location:: 303 Uris Hall
Enrollment:: 45 students (45 max) as of January 9, 2026 / Full
Notes:: Some weeks will have Friday field trips
Instructor:: Matthew I Palmer

Spring 2026 Ecology, Evolution and Environmental Biology GU4192
INTRO TO LANDSCAPE ANALYSIS

Section 001

Call Number:: 13250
Points:: 3
Day/Time:: M 8:40am-11:25am
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 20 students (25 max) as of January 9, 2026
Instructors:: Marcia N Macedo and Jay M Schoen

Spring 2026 Ecology, Evolution and Environmental Biology GU4201
ECO, BEHAVIOR & CONSERVATION OF MAMMALS

Section 001

Call Number:: 13254
Points:: 3
Day/Time:: R 6:10pm-8:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 12 students (12 max) as of January 9, 2026 / Full
Instructor:: Scott Lapoint

Spring 2026 Ecology, Evolution and Environmental Biology GU4340
HUMAN ADAPTATION

Section 001

Call Number:: 16846
Points:: 3
Day/Time:: TR 11:40am-12:55pm
Location:: 608 Schermerhorn Hall [SCH]
Enrollment:: 13 students (18 max) as of January 9, 2026
Instructor:: Volney K Friedrich

Spring 2026 Ecology, Evolution and Environmental Biology GU4670
Introduction to Geographical Information

Section 001

Call Number:: 13258
Points:: 3
Day/Time:: R 12:10pm-2:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 36 students (34 max) as of January 9, 2026 / Full
Instructor:: Viorel Popescu

Spring 2026 Ecology, Evolution and Environmental Biology GR5851
THESIS DEVELOPMENT SEMINAR

Section 001

Call Number:: 13259
Points:: 3
Day/Time:: W 10:10am-12:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 8 students (20 max) as of January 9, 2026
Instructor:: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology GR6110
FUNDAMENTALS OF EVOLUTION

Section 001

Call Number:: 13260
Points:: 3
Day/Time:: TR 8:10am-10:00am
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 22 students (30 max) as of January 9, 2026
Instructor:: Kira Delmore

Spring 2026 Ecology, Evolution and Environmental Biology GR6300
RESEARCH SEMINAR

Section 001

Call Number:: 13261
Points:: 1
Day/Time:: T 4:00pm-5:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Day/Time:: R 2:30pm-3:30pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 27 students (100 max) as of January 9, 2026
Instructor:: Maria Diuk-Wasser

Spring 2026 Ecology, Evolution and Environmental Biology GR6905
GRAD SEM IN CONSERVATION BIOL

Section 001

Call Number:: 13262
Points:: 3
Day/Time:: T 10:10am-12:00pm
Location:: 1015 EXT Schermerhorn Hall [SCH]
Enrollment:: 15 students (25 max) as of January 9, 2026
Instructor:: Bekka Brodie

Spring 2026 Ecology, Evolution and Environmental Biology GR9509
DIRECTED READINGS

Section 001

Call Number:: 18063
Points:: 1-6
Enrollment:: 1 student (2 max) as of January 9, 2026
Instructor:: Jill S Shapiro

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.

Spring 2026 Department: Ecology, Evolution, and Environmental Biology

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BEHAVIOR BIOL-LIVING PRIM

Bio Interactions-Changing

EBHS SENIOR THESIS SEMINA

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

NOTE: Students, Instructors, and Staff should use Vergil Course Search at vergil.columbia.edu/vergil/ for more detailed, accurate, and timely course information.

Spring 2026 Department: Ecology, Evolution, and Environmental Biology

For easier printing select theplain text version of this department listingand be sure to print using LANDSCAPE mode

BEHAVIOR BIOL-LIVING PRIM

Bio Interactions-Changing

EBHS SENIOR THESIS SEMINA

PRINCIPLES & APPS MOD DNA

Intermed Stats-Ecology &

INTRO TO CONSERVATION GEN

INTRO TO LANDSCAPE ANALYS

ECO, BEHAVIOR & CONSERVAT

Introduction to GIS

THESIS DEVELOPMENT SEMINA

GRAD SEM IN CONSERVATION

For easier printing select the
plain text version of this department listing
and be sure to print using LANDSCAPE mode